Physiology Coursework


(Waivers or substitutions must be approved by the director)


Cell Biology
  • BSCI 5/70143 Eukaryotic Cell Biology (3 cr) Current study of the structure and function of eukaryotic cells, including recent advances in research technology.
  • Must enroll concurrently in: BSCI 6/70144 Selected Readings in Eukaryotic Cell Biology (1 cr)
General Biochemistry

One of the following:

  • CHEM 5/70261 Principles of Biochemistry I (3 cr)Introduction to biochemical principles, including chemistry and metabolism of biochemically important compounds.
  • CHEM 5/70262 Principles of Biochemistry II (3 cr) Supramolecular and cellular biochemistry, including transcription and translation
  • BSCI 5/70142 Bioenergetics (3 cr) Respiration and photosynthesis, their origin, development, and control in living systems. Concepts are introduced from fundamental principles.
Quantitative Methods and Statistics

One of the following:

  • BSCI 6/70104 Biological Statistics (4 cr)
  • PSYC 6/71651 Statistical Inference in Psychology (3 cr)
  • BMS 78637 Analysis of BioAnthropological Data I (5 cr)
  • BMS 78638 Analysis of BioAnthropological Data II (3 cr)
  • Equivalent course approved by Director
  • BMS 6/70291 Introduction to Biomedical Sciences (1 cr)
Professional Development
  • BMS 6/71000 Responsible Conduct of Research (1 cr)

Physiology Core

Basic Physiology

Student must complete one of the following:

  • BMS 6/70449 & 6/70450 Medical Physiology I & II (7 cr)Biophysical and biochemical concepts of integrative organ system physiology in the human.
  • BSCI 5/70433 & 5/70434 Mammalian Physiology I & II (6 cr)
  • BSCI 5/70444 & 5/70445 Mammalian Physiology Laboratory I & II (2 cr) The internal environment, metabolism and energy balance, temperature regulation, nervous system and special senses, the cardiovascular, renal, respiratory and digestive systems.

Concentration Electives

Student must complete two courses from either option (a) or (b):

a. Cardiopulmonary Physiology
  • BMS 6/70451 Microcirculation (2 cr)Theory and practical application of current techniques to evaluate fluid and solute exchange across the microvascular wall.
  • BMS 6/70452 Pulmonary Physiology (2 cr) Relationship between structure and function of the lung with focus on: mechanics, circulation, gas exchange, gas transport, acid/base regulation, host-defense, ventilation-perfusion.
  • BMS 6/70429 Molecular and Cellular Neuroscience (4 cr) The relation of aspects of the neurosciences to the fundamental properties of nervous tissue, establishing a firm base in experimental neurobiology.
b. Exercise Physiology
  • ELS 6/75081 Energy Metabolism and Body Composition (3 cr) Measurement of metabolic response to exercise. Topics include ergometry, spirometry, energy expenditure, body composition, and performance correlates of strength, power, and endurance.
  • ELS 6/75082 Cardio-Respiratory Function During Exercise (3 cr) Measurement of the cardiovascular-respiratory response to exercise. Includes resting spirometry, lung function during exercise, electrocardiography, blood pressure, PWC testing and exercise prescription.

General Electives

May include courses listed above not selected, other pertinent coursework at a consortial institution not otherwise selected, and/or the following as appropriate to the student's interests and objectives:

  • BMS 6/70550 & 6/70551 Medical Pharmacology I & II (6 cr)
  • BSCI 6/70461 Neurochemistry (2 cr)
  • BSCI 5/70174 Immunology (2 cr)
  • ELS 75004 Biomechanics (3 cr)
  • ELS 75083 Exercise Energy Metabolism (3 cr)
  • ELS 75084 Cardiovascular-Respiratory Dynamics in Exercise (3 cr)
  • ELS 75085 Anthropometry and Body Comp. in the Exercise Sciences (3 cr)
  • ELS 75086 Muscle Function and Exercise (3 cr)
  • ELS 75087 Environmental Stress and Exercise (3 cr)


All students are required to register for seminars appropriate to their field of interest for a minimum of three semesters.

Additional coursework

May be required by the student's guidance committee in order to meet Program or Departmental requirements.


The teaching of laboratories and, as appropriate, lecture courses is considered important in the training and development of doctoral candidates. Students should have this experience during their graduate career, including those on non-teaching scholarships or research assistantships for most of their tenure.

Professional Development

Candidates are expected to engage, to the extent possible, in other activities beneficial to their professional development. These include membership in professional organizations; attendance at meetings, and when appropriate, presentation of research results; familiarity with the relevant literature; and other activities available during their graduate career.